Rowland,
You've got me worried now

and I have a few questions for you at the end of my post.
My observations :-
Video 1, Movement you see on bearings and seal is perfectly normal.
Video 2, Lab seal may have been designed with a clearance fit on outside, without the oring, you can see it turn slightly. Was probably to allow for greater expansion of the alloy.
Previously, I was not sure if the oring was stopping the crank from fully seating but without it, the crank is still not seating fully (IE, bearings can still be rocked) which may be due to an interference bearing fit in the cases. In fact I've taken the left 'c' ring out and I can slip a 0.5mm feeler under the bearing, but I have the oring fitted. So I now believe the crank bearings do not fully seat until the top case is bolted on, (with or without the oring). I can also rock some of the bearings slightly with the crank placed in the lower case. When you spin the crank it may be imperceptibly rocking in the case and moving the bearing outers, as in the videos
Video 3, Fitting the new oring makes things seem worse. Its groove is only 2.6mm deep, there is a lot of rubber to be compressed which the weight of the crank can't do. The crank is able to rock even more. Try giving each bearing a light tap to better seat them and see if things improve.
On old or reused cranks the oring is already squashed, /deformed and allows the crank to seat better, the bearings will also have bedded in, maybe why this wasn't noticed before on rebuilds.
A bit of a ramble (as usual for me

) but I've tried to stick to the main points. I'm sure both our cranks will be just fine when the top case gets bolted up.
Now, with my crank correctly located on the right c ring I have a gap of approx 0.7mm between the left bearing and c ring. Seems a lot. Do you have same. My whole crank seems biased to the right, based on case clearances and centre bearing positions. All the crank shims seem to be in place. Are you able to measure the flywheel gaps across the rods and centre bearings on yours. I have 8.9mm on the rods and 50.5mm on centre.
Tried a dry fit of top case, will close to about 1/16'' gap, not happy about flipping the cases to put the bottom bolts in as knock pins could displace. Will maybe support on trestle and bolt from bottom. Cases might close up better if 2 seals on left of gearbox are pushed in after assembly.
Are you intending to apply sealant with shafts in place, be easier without, but threebond I'm using advises assembly within 1 minute.
Like yourself, I rebuilt these engines, twice, 45+ years ago without issues. Are we just more cautious or experienced now ?
Brian, we'll get there.